TOEFL Reading Practice Test #42

TOEFL Reading Practice Test #42 - The Purpose of Forked Tongues in Snakes

TOEFL Reading Practice Test #42 - The Purpose of Forked Tongues in Snakes

The forked tongue of snakes has attracted attention for thousands of years, leading to a wide range of explanations. In many cultures, it has symbolized deceit or evil. Aristotle was the first person known to raise the question of its purpose, suggesting that the fork might intensify the sense of taste by doubling the sensation. By the early twentieth century, most scientists believed the tongue functioned as a tactile organ, used to probe the ground in a manner similar to how a blind person might use a cane.

In 1920, Browman introduced a theory that was widely accepted for decades. He proposed that when a snake draws its tongue back into its mouth, the tips of the fork enter two openings in the roof of the mouth, allowing chemical substances to reach a pair of vomeronasal organs (VNO). These organs, which are particularly developed in snakes, lizards, and many mammals, are part of a secondary olfactory system believed to detect pheromones—chemical messages that animals secrete to signal others of their species. According to Browman’s theory, the tongue gathers these chemical cues from the environment and transports them to the VNO. However, X-ray films taken in the 1980s revealed this was incorrect. When the tongue retracts, it enters a protective sheath and the tips do not contact the VNO openings. Instead, chemical particles are transferred to pads at the base of the mouth, and when the snake closes its mouth, the pads press the particles against the VNO openings.

If the tongue is not forked to fit into the VNO, then what might its shape accomplish? A researcher named Schwenk proposed a new hypothesis based on findings from behavior, ecology, sensory physiology, and neuroanatomy. He suggested that the forked tongue enables the snake to sense chemical cues at two separate points simultaneously, helping it detect slight differences in an odor trail. This capacity makes it easier for the animal to locate the edges of a chemical trail and follow it accurately—an ability that is important both for finding prey and locating mates.

This form of spatial chemical detection resembles other systems in which perception depends on input from two separated organs. For example, the brain determines the source of sounds by comparing what each ear hears. Likewise, using two eyes enables depth perception through stereoscopic vision.

Several pieces of evidence support the idea that forked tongues evolved to serve as chemical edge detectors that improve the tracking of odor trails. First, snakes and lizards spread the tines of their tongues apart when collecting odor particles, then bring them together when pulling the tongue back in. The farther apart the tines are when sampling, the better the animal can detect variations across the odor field. Second, lizard species that travel over wide areas in search of food generally have forked tongues, while those that do not forage widely usually lack this feature. Third, forked tongues have appeared independently in at least two separate reptile groups, suggesting that this trait has adaptive value. Finally, in snakes, each tine of the tongue is connected to a specific part of the opposite side of the brain, and those brain regions are connected to each other. This is similar to the arrangement found in the auditory centers of mammals and birds, where signals from both ears are compared to determine sound direction.

Other types of animals have also evolved paired chemical-sensing structures to locate food or mates. For example, male gypsy moths use their large, sensitive antennae to detect the pheromones of females from far away. Similarly, the ant nest beetle has spoon-like antennae extending from either side of its head that help it follow the scent trails of ants, which it preys upon.

TOEFL Reading Practice Test #42 - The Purpose of Forked Tongues in Snakes

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TOEFL Reading Practice Test #42 - The Purpose of Forked Tongues in Snakes

TOEFL Reading Practice Test #42 - The Purpose of Forked Tongues in Snakes

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1 / 10

1. The word “secrete” in the passage is closest in meaning to:

2 / 10

2. According to paragraph 2, what was discovered as a result of X-ray film studies of snakes and lizards?

3 / 10

3. The word “simultaneously” in the passage is closest in meaning to:

4 / 10

4. According to Schwenk’s hypothesis as presented in paragraphs 3 and 4, what is the advantage of a snake’s forked tongue?

5 / 10

5. According to paragraph 5, snakes and lizards are greatly aided in following odor trails by:

6 / 10

6. Which of the following is presented in paragraph 5 as evidence supporting the hypothesis that forked tongues evolved to enhance the ability to follow odor trails?

7 / 10

7. It can be inferred from paragraph 5 that the tines of the tongue in snakes are connected to the nervous system in such a way that:

8 / 10

8. Why does the author talk about “gypsy moths” and the “ant nest beetle”?

9 / 10

9. Directions:
Look at the four squares [■] that indicate where the following sentence could be added to the passage.

Sentence to Insert:
"In other words, lizard tongues are forked precisely in cases where being able to follow odor trails is an advantage."

Excerpt:
Several kinds of evidence support the hypothesis that forked tongues evolved as chemosensory edge detectors to enhance the ability to follow odor trails: Snakes and lizards spread the tines of their tongue apart when they retrieve odor molecules, then draw the tines together when retracting the tongue. The greater the distance between sampling points, the better the animals sample differences within an odor trail. [■][1] Lizards that forage widely have forked tongues, whereas lizard species without forked tongues tend not to forage widely. [■][2] Forked tongues have evolved independently at least twice in different families of reptiles, indicating their value as an adaptation. [■][3] ln the snake nervous system, each tine of the tongue is linked to a nucleus in the other side of the brain, and the two nuclei are linked across the two hemispheres. This arrangement is similar to the anatomy of auditory centers in mammals and birds that permits the computation of differences between what one ear hears and what the other ear hears and thus mediates auditory localization. [■][4]

10 / 10

10. Directions:
An introductory sentence for a brief summary of the passage is provided below. Complete the summary by selecting the THREE answer choices that express the most important ideas in the passage. Some sentences do not belong in the summary because they express ideas that are not presented in the passage or are minor details.

Introductory Sentence:
In the 1980s Browman’s widely accepted theory of why snakes have forked tongues was proven to be untrue.

Quiz Completed!

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